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The British Admiralty first contracted for a machine-gun carrying aircraft in November 1912. The manufacturer was Vickers; the machine was the No. 18, which appeared the following year at the Olympia Aero Show as the "Destroyer." This first machine was wrecked in a crash, but the followup variant was the direct ancestor of the F.B. 5 "Gunbus." All of these machines were armed with a belt-fed Vickers.
Incidentally, what appears to be a water jacket in the photo referred to is in fact the cover installed on the Lewis LMG to protect the cooling fins on the barrel.
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