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The belt-fed fixed guns were given a big magazine capacity (normally at least 250 rpg, sometimes a lot more) so didn't need to be reloaded. The top-wing mounted fixed Lewis guns (which initially used 47-round mags, later 97 rounds) were a real pain to reload until the Foster mounting allowed the pilot to pull the gun downwards first. Even then it wasn't easy, due to wind resistance acting on the drum.
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Tony Williams: Military gun and ammunition website
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