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The method of firing the machine guns by pulling a lever on the gun itself was a bit of "artistic licence" on the part of movie-makers, beginning with "The Dawn Patrol" I think. Reality is that the guns were fired from the control column; problem is that if you film this from outside the cockpit you don't see anything happening. So Hollywood decided that having the pilot pull on a lever would at least make it LOOK interesting...
The only pulling on the gun involved charging the first round with the loading lever or using it to try to clear a jam.
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